Antagonistic Transcription Factors WRKY75/WRKY46 Modulate Antioxidant Defense via SNAT4‐Mediated Melatonin Biosynthesis to Enhance Drought and Heat Stress Tolerance in Poplar

Summary statement Bu et al. reveal that PagSNAT4‐mediated regulation of endogenous melatonin biosynthesis enhances drought and heat stress tolerance in Populus . This pathway is dually regulated through antagonistic transcriptional control by PagWRKY75 and PagWRKY46, which oppositely modulate PagSNAT4 expression.