Variations in spruce (Piceasp.) distribution in the Chinese Loess Plateau and surrounding areas during the Holocene

Based on published pollen data from 44 sites in the Loess Plateau and surrounding areas, we recalculate values of Picea abundance digitized from the original diagrams and construct six isopoll maps to reconstruct the distribution of spruce forest at 2000 yr intervals during the Holocene. Spruce forest gradually expanded from about 10 ka BP ago and reached its peak distribution around 8 ka BP. It retained its maximum extent between 8 ka BP and 6 ka BP, broadly covering the subalpine area and stretching to the valleys of the western Loess Plateau with high densities and even reaching the edges of the adjacent desert regions. After 6 ka BP, spruce forest began to retreat to higher elevations. This trend was accelerated after 4 ka BP. After 2 ka BP, it disappeared completely from the Loess Plateau and most of its surrounding areas. Precipitation strongly dominated by the Asian summer monsoon was the primary controlling factor in the distribution of spruce before 2 ka BP. After about 2 ka BP, the increasing intensity of human activity became the primary factor in the disappearance of spruce in the Loess Plateau, resulting in an enormous impact on the present landscape.