Communication of queen's presence. Inhibitory effect of queens on queen rearing. Control of worker ovary development. Laying workers. Stimulatory effect of queen pheromone. Brood pheromone: recognition and stimulatory effect. Comb pheromone: recognition and stimulatory effect. Regulation of drone population. Mating pheromones. Nest and nestmate recognition. Trail and deterrent pheromones. Attraction. Nasonov pheromone. Alarm and aggression pheromones. Bumblebees: colony and foraging pheromones. Bumblebees: mating behaviour.